About the Authors

Raul A. Cernadas

Affiliations Department of Plant Pathology and Microbiology, Iowa State University, Ames, Iowa, United States of America, Department of Plant Pathology and Plant-Microbe Biology, Cornell University, Ithaca, New York, United States of America

Erin L. Doyle

Address Current address: Department of Biology, Doane College, Crete, Nebraska, United States of America.

Affiliations Department of Plant Pathology and Microbiology, Iowa State University, Ames, Iowa, United States of America, Bioinformatics and Computational Biology Graduate Program, Iowa State University, Ames, Iowa, United States of America

David O. Niño-Liu

Address Current address: Monsanto Company, St. Louis, Missouri, United States of America.

Affiliation Department of Plant Pathology and Microbiology, Iowa State University, Ames, Iowa, United States of America

Katherine E. Wilkins

Affiliation Department of Plant Pathology and Plant-Microbe Biology, Cornell University, Ithaca, New York, United States of America

Timothy Bancroft

Address Current address: Health Economics and Outcomes Research, OptumInsight, Eden Prairie, Minnesota, United States of America.

Affiliation Department of Statistics, Iowa State University, Ames, Iowa, United States of America

Li Wang

Affiliations Department of Plant Pathology and Microbiology, Iowa State University, Ames, Iowa, United States of America, Department of Plant Pathology and Plant-Microbe Biology, Cornell University, Ithaca, New York, United States of America

Clarice L. Schmidt

Affiliation Department of Plant Pathology and Microbiology, Iowa State University, Ames, Iowa, United States of America

Rico Caldo

Address Current address: Monsanto Company, St. Louis, Missouri, United States of America.

Affiliation Department of Plant Pathology and Microbiology, Iowa State University, Ames, Iowa, United States of America

Bing Yang

Affiliation Genetics Development and Cell Biology, Iowa State University, Ames, Iowa, United States of America

Frank F. White

Affiliation Department of Plant Pathology, Kansas State University, Manhattan, Kansas, United States of America

Dan Nettleton

Affiliation Department of Statistics, Iowa State University, Ames, Iowa, United States of America

Roger P. Wise

Affiliations Department of Plant Pathology and Microbiology, Iowa State University, Ames, Iowa, United States of America, Corn Insects and Crop Genetics Research, USDA-ARS, Iowa State University, Ames, Iowa, United States of America

Adam J. Bogdanove

ajb7@cornell.edu

Affiliations Department of Plant Pathology and Microbiology, Iowa State University, Ames, Iowa, United States of America, Department of Plant Pathology and Plant-Microbe Biology, Cornell University, Ithaca, New York, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: RAC DONL KEW LW CLS RC FFW DN RPW AJB. Performed the experiments: RAC DONL KEW LW CLS RC BY. Analyzed the data: RAC ELD DONL KEW TB LW RC DN AJB. Contributed reagents/materials/analysis tools: CLS BY FFW RPW. Wrote the paper: RAC ELD KEW DN RPW AJB.