About the Authors

Christopher J. Graves

Affiliation University of Pennsylvania, Philadelphia, Pennsylvania, United States of America

Vera I. D. Ros

Affiliation University of Pennsylvania, Philadelphia, Pennsylvania, United States of America

Brian Stevenson

Affiliation University of Kentucky, Lexington, Kentucky, United States of America

Paul D. Sniegowski

Affiliation University of Pennsylvania, Philadelphia, Pennsylvania, United States of America

Dustin Brisson

dbrisson@sas.upenn.edu

Affiliation University of Pennsylvania, Philadelphia, Pennsylvania, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: DB CJG. Performed the experiments: CJG VIDR DB. Analyzed the data: CJG DB PDS. Contributed reagents/materials/analysis tools: BS. Wrote the paper: CJG PDS DB.