About the Authors

Joseph Hughes

Affiliation Medical Research Council-University of Glasgow Centre for Virus Research, Institute of Infection, Inflammation and Immunity, College of Medical, Veterinary and Life Sciences, University of Glasgow, Glasgow, United Kingdom

Richard C. Allen

Affiliation Disease Dynamics Unit, Department of Veterinary Medicine, University of Cambridge, Cambridge, United Kingdom

Marc Baguelin

Affiliations Immunisation, Hepatitis and Blood Safety Department, Health Protection Agency, London, United Kingdom, Centre for Mathematical Modelling of Infectious Diseases, London School of Hygiene and Tropical Medicine, London, United Kingdom

Katie Hampson

Affiliation Boyd Orr Centre for Population and Ecosystem Health, Institute for Biodiversity, Animal Health and Comparative Medicine, University of Glasgow, Glasgow, United Kingdom

Gregory J. Baillie

Affiliation Wellcome Trust Sanger Institute, Wellcome Trust Genome Campus, Hinxton, Cambridge, United Kingdom

Debra Elton

Affiliation Animal Health Trust, Centre for Preventive Medicine, Lanwades Park, Newmarket, United Kingdom

J. Richard Newton

Affiliation Animal Health Trust, Centre for Preventive Medicine, Lanwades Park, Newmarket, United Kingdom

Paul Kellam

Affiliation Wellcome Trust Sanger Institute, Wellcome Trust Genome Campus, Hinxton, Cambridge, United Kingdom

James L. N. Wood

Affiliation Disease Dynamics Unit, Department of Veterinary Medicine, University of Cambridge, Cambridge, United Kingdom

Edward C. Holmes

Affiliations Center for Infectious Disease Dynamics, Department of Biology, The Pennsylvania State University, University Park, Pennsylvania, United States of America, Fogarty International Center, National Institute of Health, Bethesda, Maryland, United States of America

Pablo R. Murcia

Pablo.Murcia@Glasgow.ac.uk

Affiliations Medical Research Council-University of Glasgow Centre for Virus Research, Institute of Infection, Inflammation and Immunity, College of Medical, Veterinary and Life Sciences, University of Glasgow, Glasgow, United Kingdom, Disease Dynamics Unit, Department of Veterinary Medicine, University of Cambridge, Cambridge, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: PRM. Performed the experiments: RCA. Analyzed the data: JH PRM ECH MB KH. Contributed reagents/materials/analysis tools: DE JRN PK GJB. Wrote the paper: JH PRM ECH JLNW MB KH.