About the Authors

Roman Biek

Contributed equally to this work with: Roman Biek, Anthony O'Hare

Affiliation Boyd Orr Centre for Population and Ecosystem Health and Institute for Biodiversity, Animal Health and Comparative Medicine, University of Glasgow, Glasgow, United Kingdom

Anthony O'Hare

Contributed equally to this work with: Roman Biek, Anthony O'Hare

Affiliation Boyd Orr Centre for Population and Ecosystem Health and Institute for Biodiversity, Animal Health and Comparative Medicine, University of Glasgow, Glasgow, United Kingdom

David Wright

Affiliations Veterinary Sciences Division, Agri-Food and Biosciences Institute, Stormont, Belfast, Northern Ireland, United Kingdom, School of Biological Sciences, Queen's University Belfast, Belfast, Northern Ireland, United Kingdom

Tom Mallon

Affiliation Veterinary Sciences Division, Agri-Food and Biosciences Institute, Stormont, Belfast, Northern Ireland, United Kingdom

Carl McCormick

Affiliation Veterinary Sciences Division, Agri-Food and Biosciences Institute, Stormont, Belfast, Northern Ireland, United Kingdom

Richard J. Orton

Affiliation Boyd Orr Centre for Population and Ecosystem Health and Institute for Biodiversity, Animal Health and Comparative Medicine, University of Glasgow, Glasgow, United Kingdom

Stanley McDowell

Affiliation Veterinary Sciences Division, Agri-Food and Biosciences Institute, Stormont, Belfast, Northern Ireland, United Kingdom

Hannah Trewby

Affiliation Boyd Orr Centre for Population and Ecosystem Health and Institute for Biodiversity, Animal Health and Comparative Medicine, University of Glasgow, Glasgow, United Kingdom

Robin A. Skuce

Affiliation Veterinary Sciences Division, Agri-Food and Biosciences Institute, Stormont, Belfast, Northern Ireland, United Kingdom

Rowland R. Kao

rowland.kao@glasgow.ac.uk

Affiliation Boyd Orr Centre for Population and Ecosystem Health and Institute for Biodiversity, Animal Health and Comparative Medicine, University of Glasgow, Glasgow, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: RB AO TM SM RAS RRK. Performed the experiments: SM CM. Analyzed the data: RB AO DW RJO HT RRK. Wrote the paper: RB AOH RRK.