About the Authors

Peter J. Kerr

Affiliation CSIRO Ecosystem Sciences, Canberra, Australian Capital Territory, Australia

Elodie Ghedin

Affiliation Center for Vaccine Research, Department of Computational and Systems Biology, University of Pittsburgh School of Medicine, Pittsburgh, Pennsylvania, United States of America

Jay V. DePasse

Affiliation Center for Vaccine Research, Department of Computational and Systems Biology, University of Pittsburgh School of Medicine, Pittsburgh, Pennsylvania, United States of America

Adam Fitch

Affiliation Center for Vaccine Research, Department of Computational and Systems Biology, University of Pittsburgh School of Medicine, Pittsburgh, Pennsylvania, United States of America

Isabella M. Cattadori

Affiliation Center for Infectious Disease Dynamics, Department of Biology, The Pennsylvania State University, University Park, Pennsylvania, United States of America

Peter J. Hudson

Affiliation Center for Infectious Disease Dynamics, Department of Biology, The Pennsylvania State University, University Park, Pennsylvania, United States of America

David C. Tscharke

Affiliation Research School of Biology, The Australian National University, Canberra, Australian Capital Territory, Australia

Andrew F. Read

Affiliations Center for Infectious Disease Dynamics, Department of Biology, The Pennsylvania State University, University Park, Pennsylvania, United States of America, Fogarty International Center, National Institutes of Health, Bethesda, Maryland, United States of America

Edward C. Holmes

ech15@psu.edu

Affiliations Center for Infectious Disease Dynamics, Department of Biology, The Pennsylvania State University, University Park, Pennsylvania, United States of America, Fogarty International Center, National Institutes of Health, Bethesda, Maryland, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: PJK EG IMC PJH AFR ECH. Performed the experiments: PJK EG JVD AF. Analyzed the data: PJK EG JVD AF ECH. Contributed reagents/materials/analysis tools: PJK EG JVD AF DCT. Wrote the paper: PJK EG IMC PJH DCT AFR ECH.