About the Authors

Michael D. Alpert

Affiliation Department of Microbiology and Immunobiology, Harvard Medical School, New England Primate Research Center, Southborough, Massachusetts, United States of America

Jackson D. Harvey

Affiliation Department of Microbiology and Immunobiology, Harvard Medical School, New England Primate Research Center, Southborough, Massachusetts, United States of America

W. Anderson Lauer

Affiliation Department of Microbiology and Immunobiology, Harvard Medical School, New England Primate Research Center, Southborough, Massachusetts, United States of America

R. Keith Reeves

Affiliation Immunology Division, Harvard Medical School, New England Primate Research Center, Southborough, Massachusetts, United States of America

Michael Piatak Jr.

Affiliation SAIC Frederick, National Cancer Institute at Frederick, Frederick, Maryland, United States of America

Angela Carville

Affiliation Department of Pathology, Harvard Medical School, New England Primate Research Center, Southborough, Massachusetts, United States of America

Keith G. Mansfield

Affiliation Department of Pathology, Harvard Medical School, New England Primate Research Center, Southborough, Massachusetts, United States of America

Jeffrey D. Lifson

Affiliation SAIC Frederick, National Cancer Institute at Frederick, Frederick, Maryland, United States of America

Wenjun Li

Affiliation University of Massachusetts Medical School, Worcester, Massachusetts, United States of America

Ronald C. Desrosiers

Affiliation Department of Microbiology and Immunobiology, Harvard Medical School, New England Primate Research Center, Southborough, Massachusetts, United States of America

R. Paul Johnson

Affiliations Immunology Division, Harvard Medical School, New England Primate Research Center, Southborough, Massachusetts, United States of America, Ragon Institute of Massachusetts General Hospital, MIT, and Harvard, and Infectious Disease Unit, Massachusetts General Hospital, Boston, Massachusetts, United States of America

David T. Evans

david_evans@hms.harvard.edu

Affiliation Department of Microbiology and Immunobiology, Harvard Medical School, New England Primate Research Center, Southborough, Massachusetts, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: MDA RKR RCD RPJ DTE. Performed the experiments: MDA JDH. Analyzed the data: MDA WL RPJ RCD DTE. Wrote the paper: MDA DTE. Provided technical assistance: WAL. Performed the viral load measurements: MP JDL. Performed veterinary procedures: AC KGM.