About the Authors

Justin Lessler, Derek A. T. Cummings
Department of Epidemiology, Johns Hopkins Bloomberg School of Public Health, Baltimore, Maryland, United States of America

Steven Riley
School of Public Health, Department of Infectious Disease Epidemiology, MRC Centre for Outbreak Analysis and Modelling, Imperial College, London, United Kingdom

Jonathan M. Read
Department of Epidemiology and Population Health, Institute of Infection and Global Health, Faculty of Health and Life Sciences, University of Liverpool, Liverpool, United Kingdom

Shuying Wang, Chao Qiang Jiang
Guangzhou No. 12 Hospital, Guangzhou, Guangdong, China

Huachen Zhu, Yi Guan
International Institute of Infection and Immunity, Shantou University Medical College, Shantou, Guangdong, China

Huachen Zhu, Yi Guan
Department of Microbiology, The University of Hong Kong, Hong Kong SAR, China

Gavin J. D. Smith
Laboratory of Virus Evolution, Program in Emerging Infectious Diseases, Duke-NUS Graduate Medical School, Singapore, Singapore

Corresponding Author

s.riley@imperial.ac.uk

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: SR DATC JMR GJDS CQJ YG. Performed the experiments: SW HZ. Analyzed the data: SR DATC JMR. Wrote the paper: JL SR JMR DATC.