About the Authors

Katharine J. Bar, Shilpa S. Iyer, Hui Li, Shuyi Wang, Beatrice H. Hahn, George M. Shaw
Perelman School of Medicine, University of Pennsylvania, Philadelphia, Pennsylvania, United States of America

Chun-yen Tsao, Mattia Bonsignori, Xi Chen, Kwan-Ki Hwang, David C. Montefiori, Hua-Xin Liao, Jeffrey W. Pavlicek, Feng Gao, Barton F. Haynes
Duke University School of Medicine, Durham, North Carolina, United States of America

Julie M. Decker, Sarah Sterrett
University of Alabama at Birmingham, Birmingham, Alabama, United States of America

Yongping Yang, Ivelin Georgiev, Jason S. McLellan, Peter D. Kwong
Vaccine Research Center, National Institute of Allergy and Infectious Disease, National Institutes of Health, Bethesda, Maryland, United States of America

Peter Hraber, William Fischer, Alan S. Perelson, Bette T. Korber
Los Alamos National Laboratory, Los Alamos, New Mexico, United States of America

Brandon F. Keele
SAIC-Frederick Inc, National Cancer Institute, Frederick, Maryland, United States of America

Vitaly V. Ganusov
University of Tennessee, Knoxville, Tennessee, United States of America

Corresponding Author

shawg@upenn.edu

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: KJB CT MB WF VVG ASP BTK JSM PDK GMS. Performed the experiments: KJB CT SSI JMD YY MB XC KH HXL HL PH WF SW SS VVG JWP. Analyzed the data: KJB DCM HXL HL BFK VVG ASP BTK IG JSM FG BFH BHH PDK GMS. Wrote the paper: KJB VVG ASP BTK IG BFH BHH PDK GMS.