About the Authors

Bradley S. Barrett

Affiliation Department of Medicine, University of Colorado Denver, Aurora, Colorado, United States of America

Diana S. Smith

Affiliation Department of Medicine, University of Colorado Denver, Aurora, Colorado, United States of America

Sam X. Li

Affiliation Department of Microbiology, University of Colorado Denver, Aurora, Colorado, United States of America

Kejun Guo

Affiliation Department of Medicine, University of Colorado Denver, Aurora, Colorado, United States of America

Kim J. Hasenkrug

Affiliation Rocky Mountain Laboratories, National Institutes for Allergy and Infectious Diseases, National Institutes of Health (NIH), Hamilton, Montana, United States of America

Mario L. Santiago

mario.santiago@ucdenver.edu

Affiliations Department of Medicine, University of Colorado Denver, Aurora, Colorado, United States of America, Department of Microbiology, University of Colorado Denver, Aurora, Colorado, United States of America, Department of Immunology, University of Colorado Denver, Aurora, Colorado, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: MLS BSB KJH. Performed the experiments: BSB MLS DSS SXL KG. Analyzed the data: MLS BSB KJH DSS SXL KG. Wrote the paper: MLS BSB KJH.