About the Authors

Elizabeth Redman, Fiona Whitelaw
Institute of Infection, Immunity and Inflammation, College of Medical, Veterinary and Life Sciences, University of Glasgow, Glasgow, United Kingdom

Neil Sargison, Frank Jackson, Alison Morrison, David Jon Bartley
Moredun Research Institute, Pentlands Science Park, Midlothian, United Kingdom

John Stuart Gilleard
Department of Comparative Biology and Experimental Medicine, Faculty of Veterinary Medicine, University of Calgary, Calgary, Alberta, Canada

Corresponding Author


Competing Interests

I have read the journal's policy and have the following conflicts. The study received some funding support from Pfizer Animal Health PLC. This does not alter our adherence to all the PLoS Pathogens policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: JSG NS ER. Performed the experiments: NS ER FJ AAM DJB FW. Analyzed the data: NS ER DJB. Wrote the paper: NS ER JSG. Backcrossing and animal work: NS ER FJ AAM DJB. Genotyping: ER NS FW. Supervision and critical reading of the manuscript: JSG DJB.