About the Authors

Jie Lu

Affiliation Department of Microbiology and Tumor Virology Program of the Abramson Cancer Center, Perelman School of Medicine at the University of Pennsylvania, Philadelphia, Pennsylvania, United States of America

Subhash C. Verma

Affiliation Department of Microbiology and Immunology, School of Medicine, University of Nevada, Reno, Nevada, United States of America

Qiliang Cai

Affiliation Department of Microbiology and Tumor Virology Program of the Abramson Cancer Center, Perelman School of Medicine at the University of Pennsylvania, Philadelphia, Pennsylvania, United States of America

Abhik Saha

Affiliation Department of Microbiology and Tumor Virology Program of the Abramson Cancer Center, Perelman School of Medicine at the University of Pennsylvania, Philadelphia, Pennsylvania, United States of America

Richard Kuo Dzeng

Affiliation Department of Microbiology and Tumor Virology Program of the Abramson Cancer Center, Perelman School of Medicine at the University of Pennsylvania, Philadelphia, Pennsylvania, United States of America

Erle S. Robertson

erle@mail.med.upenn.edu

Affiliation Department of Microbiology and Tumor Virology Program of the Abramson Cancer Center, Perelman School of Medicine at the University of Pennsylvania, Philadelphia, Pennsylvania, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: JL SCV ESR. Performed the experiments: JL QC RKD. Analyzed the data: JL SCV. Contributed reagents/materials/analysis tools: AS. Wrote the paper: JL ESR.