About the Authors

Toshihiro Ito

Affiliations Department of Pathology, University of Michigan Medical School, Ann Arbor, Michigan, United States of America, Department of Pathology and Experimental Medicine, Graduate School of Medicine, Dentistry and Pharmaceutical Sciences, Okayama University, Okayama, Japan

Ronald M. Allen

Affiliation Department of Pathology, University of Michigan Medical School, Ann Arbor, Michigan, United States of America

William F. Carson IV

Affiliation Department of Pathology, University of Michigan Medical School, Ann Arbor, Michigan, United States of America

Matthew Schaller

Affiliation Department of Pathology, University of Michigan Medical School, Ann Arbor, Michigan, United States of America

Karen A. Cavassani

Affiliation Department of Pathology, University of Michigan Medical School, Ann Arbor, Michigan, United States of America

Cory M. Hogaboam

Affiliation Department of Pathology, University of Michigan Medical School, Ann Arbor, Michigan, United States of America

Nicholas W. Lukacs

Affiliation Department of Pathology, University of Michigan Medical School, Ann Arbor, Michigan, United States of America

Akihiro Matsukawa

Affiliation Department of Pathology and Experimental Medicine, Graduate School of Medicine, Dentistry and Pharmaceutical Sciences, Okayama University, Okayama, Japan

Steven L. Kunkel

slkunkel@umich.edu

Affiliation Department of Pathology, University of Michigan Medical School, Ann Arbor, Michigan, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: TI KAC CMH NWL AM SLK. Performed the experiments: TI RMA. Analyzed the data: TI. Contributed reagents/materials/analysis tools: TI WFC MS. Wrote the paper: TI SLK.