About the Authors

Xiaojun Ning

Affiliation Department of Microbiology and Immunology, The Penn State University College of Medicine, Hershey, Pennsylvania, United States of America

David Nguyen

Address Current address: Department of Urology and Jonsson Comprehensive Cancer Center, David Geffen School of Medicine, University of California Los Angeles, Los Angeles, California, United States of America

Affiliation Department of Microbiology and Immunology, The Penn State University College of Medicine, Hershey, Pennsylvania, United States of America

Laura Mentzer

Affiliation Department of Microbiology and Immunology, The Penn State University College of Medicine, Hershey, Pennsylvania, United States of America

Christina Adams

Affiliation Department of Microbiology and Immunology, The Penn State University College of Medicine, Hershey, Pennsylvania, United States of America

Hyunwook Lee

Affiliations Department of Microbiology and Immunology, The Penn State University College of Medicine, Hershey, Pennsylvania, United States of America, Division of Infectious Diseases, Department of Medicine, The Penn State University College of Medicine, Hershey, Pennsylvania, United States of America

Robert Ashley

Affiliations Department of Microbiology and Immunology, The Penn State University College of Medicine, Hershey, Pennsylvania, United States of America, Division of Infectious Diseases, Department of Medicine, The Penn State University College of Medicine, Hershey, Pennsylvania, United States of America

Susan Hafenstein

Affiliations Department of Microbiology and Immunology, The Penn State University College of Medicine, Hershey, Pennsylvania, United States of America, Division of Infectious Diseases, Department of Medicine, The Penn State University College of Medicine, Hershey, Pennsylvania, United States of America

Jianming Hu

juh13@psu.edu

Affiliation Department of Microbiology and Immunology, The Penn State University College of Medicine, Hershey, Pennsylvania, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: XN JH. Performed the experiments: XN DN LM CA HL RA. Analyzed the data: XN DN LM CA HL RA SH JH. Wrote the paper: XN HL SH JH.