About the Authors

Santiago Castillo-Ramírez

Affiliation Department of Biology and Biochemistry, University of Bath, Claverton Down, Bath, United Kingdom

Simon R. Harris

Affiliation The Wellcome Trust Sanger Institute, Wellcome Trust Genome Campus, Hinxton, Cambridge, United Kingdom

Matthew T. G. Holden

Affiliation The Wellcome Trust Sanger Institute, Wellcome Trust Genome Campus, Hinxton, Cambridge, United Kingdom

Miao He

Affiliation The Wellcome Trust Sanger Institute, Wellcome Trust Genome Campus, Hinxton, Cambridge, United Kingdom

Julian Parkhill

Affiliation The Wellcome Trust Sanger Institute, Wellcome Trust Genome Campus, Hinxton, Cambridge, United Kingdom

Stephen D. Bentley

Affiliation The Wellcome Trust Sanger Institute, Wellcome Trust Genome Campus, Hinxton, Cambridge, United Kingdom

Edward J. Feil

e.feil@bath.ac.uk

Affiliation Department of Biology and Biochemistry, University of Bath, Claverton Down, Bath, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: SCR SH MTGH MH JP SDB EJF. Performed the experiments: SH MTGH MH SDB. Analyzed the data: SCR SH MTGH JP MH EJF. Contributed reagents/materials/analysis tools: SCR SH JP SDB. Wrote the paper: SCR EJF.