About the Authors

Alison Crawford, Jill Marie Angelosanto, Shawn D. Blackburn, E. John Wherry
Department of Microbiology and Institute for Immunology, University of Pennsylvania School of Medicine, Philadelphia, Pennsylvania, United States of America

Kim Lynn Nadwodny
GlaxoSmithKline, Department of Safety Assessment, Immunologic Toxicology, King of Prussia, Pennsylvania, United States of America

Corresponding Author


Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: AC EJW. Performed the experiments: AC JMA SDB KLN. Analyzed the data: AC. Wrote the paper: AC EJW.