About the Authors

Alison Crawford

Affiliation Department of Microbiology and Institute for Immunology, University of Pennsylvania School of Medicine, Philadelphia, Pennsylvania, United States of America

Jill Marie Angelosanto

Affiliation Department of Microbiology and Institute for Immunology, University of Pennsylvania School of Medicine, Philadelphia, Pennsylvania, United States of America

Kim Lynn Nadwodny

Affiliation GlaxoSmithKline, Department of Safety Assessment, Immunologic Toxicology, King of Prussia, Pennsylvania, United States of America

Shawn D. Blackburn

Affiliation Department of Microbiology and Institute for Immunology, University of Pennsylvania School of Medicine, Philadelphia, Pennsylvania, United States of America

E. John Wherry

Wherry@mail.med.upenn.edu

Affiliation Department of Microbiology and Institute for Immunology, University of Pennsylvania School of Medicine, Philadelphia, Pennsylvania, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: AC EJW. Performed the experiments: AC JMA SDB KLN. Analyzed the data: AC. Wrote the paper: AC EJW.