About the Authors

Martha I. Nelson, Yi Tan, Cécile Viboud, Andrew Rambaut, Edward C. Holmes
Division of International Epidemiology and Population Studies, Fogarty International Center, National Institutes of Health, Bethesda, Maryland, United States of America

Philippe Lemey
Department of Microbiology and Immunology, Katholieke Universiteit Leuven, Leuven, Belgium

Amy Vincent
Virus and Prion Diseases of Livestock Research Unit, National Animal Disease Center, USDA-ARS, Ames, Iowa, United States of America

Tommy Tsan-Yuk Lam, Edward C. Holmes
Department of Biology, The Pennsylvania State University, University Park, Pennsylvania, United States of America

Susan Detmer, Marie Gramer
The University of Minnesota Veterinary Diagnostic Laboratory, St. Paul, Minnesota, United States of America

Marc A. Suchard
Departments of Biomathematics and Human Genetics, David Geffen School of Medicine at UCLA, and Department of Biostatistics, UCLA School of Public Health, Los Angeles, California, United States of America

Andrew Rambaut
Institute of Evolutionary Biology, University of Edinburgh, Ashworth Laboratories, Edinburgh, United Kingdom

Corresponding Author

nelsonma@mail.nih.gov

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: MIN PL MAS AR ECH. Performed the experiments: AV SD MG. Analyzed the data: MIN PL YT TTYL MAS CV AR ECH. Wrote the paper: MIN PL AV CV MAS AR ECH MG.