About the Authors

Craig B. Wilen, John C. Tilton, Scott A. Sherrill-Mix, Sean C. Patro, Andrea P. O. Jordan, James A. Hoxie, Frederic D. Bushman, Robert W. Doms
Department of Microbiology, University of Pennsylvania School of Medicine, Philadelphia, Pennsylvania, United States of America

Jianbin Wang, Jeffrey C. Miller, Kenneth A. Kim, Edward J. Rebar, Gary Lee, Joshua Kahn, Philip D. Gregory, Michael C. Holmes
Sangamo BioSciences, Richmond, California, United States of America

Anthony J. Secreto, Gwenn A. Danet-Desnoyers
Department of Medicine, University of Pennsylvania School of Medicine, Philadelphia, Pennsylvania, United States of America

Pyone P. Aye, Bruce A. Bunnell, Andrew A. Lackner
Divisions of Regenerative Medicine and Comparative Pathology, Tulane National Primate Research Center, Tulane University School of Medicine, Covington, Louisiana, United States of America

James L. Riley, Carl H. June
Abramson Family Cancer Research Institute, University of Pennsylvania School of Medicine, Philadelphia, Pennsylvania, United States of America

Corresponding Author

doms@mail.med.upenn.edu

Competing Interests

A number of the authors are employed by Sangamo Biosciences. The work was funded by NIH.

Author Contributions

Conceived and designed the experiments: CBW JW JCT EJR SCP JAH GADD JLR PDG CHJ MCH RWD. Performed the experiments: CBW JW KAK SCP AJS APOJ. Analyzed the data: CBW JW JCM SCP. Contributed reagents/materials/analysis tools: SASM GL JK PPA BAB AAL FDB. Wrote the paper: CBW RWD.