About the Authors

Scott A. C. Godfrey

Affiliation Faculty of Health and Life Sciences, University of the West of England, Bristol, United Kingdom

Helen C. Lovell

Affiliation Faculty of Health and Life Sciences, University of the West of England, Bristol, United Kingdom

John W. Mansfield

Affiliation Department of Life Sciences, Imperial College London, London, United Kingdom

David S. Corry

Affiliation Faculty of Health and Life Sciences, University of the West of England, Bristol, United Kingdom

Robert W. Jackson

Affiliation School of Biological Sciences, University of Reading, Reading, United Kingdom

Dawn L. Arnold

Dawn.Arnold@uwe.ac.uk

Affiliation Faculty of Health and Life Sciences, University of the West of England, Bristol, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: SACG, JWM, HCL, RWJ, DLA. Performed the experiments: SACG, HCL, DSC, JWM. Analyzed the data: SACG, HCL, DLA. Contributed reagents/materials/analysis tools: SACG, DLA, JWM. Wrote the paper: JWM, SACG, DLA, HCL, RWJ.