About the Authors

Niina Lietzén

Affiliation Institute of Biotechnology, University of Helsinki, Helsinki, Finland

Tiina Öhman

Contributed equally to this work with: Tiina Öhman, Johanna Rintahaka

Affiliation Institute of Biotechnology, University of Helsinki, Helsinki, Finland

Johanna Rintahaka

Contributed equally to this work with: Tiina Öhman, Johanna Rintahaka

Affiliation Finnish Institute of Occupational Health, Helsinki, Finland

Ilkka Julkunen

Affiliation National Institute for Health and Welfare, Helsinki, Finland

Tero Aittokallio

Affiliation Department of Mathematics, University of Turku, Turku, Finland

Sampsa Matikainen

These authors also contributed equally to this work.

Affiliation Finnish Institute of Occupational Health, Helsinki, Finland

Tuula A. Nyman

tuula.nyman@helsinki.fi

These authors also contributed equally to this work.

Affiliation Institute of Biotechnology, University of Helsinki, Helsinki, Finland

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: SM TAN. Performed the experiments: NL TÖ JR. Analyzed the data: NL TÖ TA SM TAN. Contributed reagents/materials/analysis tools: IJ TA. Wrote the paper: NL SM TAN.