About the Authors

Steve Atkinson

Affiliation School of Molecular Medical Sciences, Centre for Biomolecular Science, University of Nottingham, Nottingham, United Kingdom

Robert J. Goldstone

Affiliation School of Molecular Medical Sciences, Centre for Biomolecular Science, University of Nottingham, Nottingham, United Kingdom

George W. P. Joshua

Affiliation Department of Infectious and Tropical Diseases, London School of Hygiene and Tropical Medicine, London, United Kingdom

Chien-Yi Chang

Affiliation School of Molecular Medical Sciences, Centre for Biomolecular Science, University of Nottingham, Nottingham, United Kingdom

Hannah L. Patrick

Affiliation School of Molecular Medical Sciences, Centre for Biomolecular Science, University of Nottingham, Nottingham, United Kingdom

Miguel Cámara

Affiliation School of Molecular Medical Sciences, Centre for Biomolecular Science, University of Nottingham, Nottingham, United Kingdom

Brendan W. Wren

Affiliation Department of Infectious and Tropical Diseases, London School of Hygiene and Tropical Medicine, London, United Kingdom

Paul Williams

paul.williams@nottingham.ac.uk

Affiliation School of Molecular Medical Sciences, Centre for Biomolecular Science, University of Nottingham, Nottingham, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: SA RJG GWPJ CYC HLP MC BWW PW. Performed the experiments: SA RJG GWPJ CYC HLP PW. Analyzed the data: SA RJG GWPJ CYC HLP MC BWW PW. Contributed reagents/materials/analysis tools: SA MC. Wrote the paper: SA GWPJ MC BWW PW.