About the Authors

Zhiyong Shao

Contributed equally to this work with: Zhiyong Shao, Yi Zhang

Affiliation Department of Genetics, Development, and Cell Biology, Iowa State University, Ames, Iowa, United States of America

Yi Zhang

Contributed equally to this work with: Zhiyong Shao, Yi Zhang

Affiliation Department of Genetics, Development, and Cell Biology, Iowa State University, Ames, Iowa, United States of America

Qi Ye

Affiliation Department of Genetics, Development, and Cell Biology, Iowa State University, Ames, Iowa, United States of America

Jenifer Neeta Saldanha

Affiliation Department of Genetics, Development, and Cell Biology, Iowa State University, Ames, Iowa, United States of America

Jo Anne Powell-Coffman

japc@iastate.edu

Affiliation Department of Genetics, Development, and Cell Biology, Iowa State University, Ames, Iowa, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: ZS YZ JAPC. Performed the experiments: ZS YZ QY JNS. Analyzed the data: ZS YZ QY JNS JAPC. Contributed reagents/materials/analysis tools: ZS YZ. Wrote the paper: ZS YZ JAPC.