About the Authors

Vladimir A. Belyi

Affiliation Simons Center for Systems Biology, Institute for Advanced Study, Princeton, New Jersey, United States of America

Arnold J. Levine

am_skalka@fccc.edu (AMS); alevine@ias.edu (AJL)

Affiliation Simons Center for Systems Biology, Institute for Advanced Study, Princeton, New Jersey, United States of America

Anna Marie Skalka

am_skalka@fccc.edu (AMS); alevine@ias.edu (AJL)

Affiliation Institute for Cancer Research, Fox Chase Cancer Center, Philadelphia, Pennsylvania, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: VAB AMS. Performed the experiments: VAB AMS. Analyzed the data: VAB AJL AMS. Contributed reagents/materials/analysis tools: AMS. Conducted the computational studies and analyses, prepared the Tables and several Figures, and wrote part of the manuscript: VAB. Contributed virological expertise, supervised the computational studies, and wrote part of the manuscript: AJL. Initiated the collaboration with VAB: AMS. Contributed virological/biological expertise, designed several Figures, and wrote part of the manuscript: AMS. Contributed biological context: AMS.