About the Authors

Ben Raymond
School of Biological Sciences, Royal Holloway University of London, Egham, Surrey, United Kingdom

Ben Raymond, Kelly L. Wyres, Samuel K. Sheppard, Michael B. Bonsall
Department of Zoology, University of Oxford, Oxford, United Kingdom

Richard J. Ellis
Molecular Pathogenesis and Genetics, Veterinary Laboratories Agency, Addlestone, United Kingdom

Corresponding Author


Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: BR RJE MBB. Performed the experiments: BR KLW. Analyzed the data: BR KLW SKS. Contributed reagents/materials/analysis tools: SKS. Wrote the paper: BR KLW SKS RJE MBB.