About the Authors

Ben Raymond

ben.raymond@rhul.ac.uk

These authors are joint first authors.

Affiliations School of Biological Sciences, Royal Holloway University of London, Egham, Surrey, United Kingdom, Department of Zoology, University of Oxford, Oxford, United Kingdom

Kelly L. Wyres

These authors are joint first authors.

Affiliation Department of Zoology, University of Oxford, Oxford, United Kingdom

Samuel K. Sheppard

Affiliation Department of Zoology, University of Oxford, Oxford, United Kingdom

Richard J. Ellis

Affiliation Molecular Pathogenesis and Genetics, Veterinary Laboratories Agency, Addlestone, United Kingdom

Michael B. Bonsall

Affiliation Department of Zoology, University of Oxford, Oxford, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: BR RJE MBB. Performed the experiments: BR KLW. Analyzed the data: BR KLW SKS. Contributed reagents/materials/analysis tools: SKS. Wrote the paper: BR KLW SKS RJE MBB.