About the Authors

Bryony A. P. Williams

Affiliations School of Biosciences, Geoffrey Pope Building, University of Exeter, Exeter, Devon, United Kingdom, Department of Botany, University of British Columbia, Vancouver, British Columbia, Canada

Catherine Elliot

Affiliation Department of Biochemistry and Biomedical Sciences, School of Life Sciences, University of Sussex, Falmer, Brighton, United Kingdom

Lena Burri

Affiliation Department of Botany, University of British Columbia, Vancouver, British Columbia, Canada

Yasutoshi Kido

Affiliation Department of Biomedical Chemistry, Graduate School of Medicine, University of Tokyo, Tokyo, Japan

Kiyoshi Kita

Affiliation Department of Biomedical Chemistry, Graduate School of Medicine, University of Tokyo, Tokyo, Japan

Anthony L. Moore

Affiliation Department of Biochemistry and Biomedical Sciences, School of Life Sciences, University of Sussex, Falmer, Brighton, United Kingdom

Patrick J. Keeling

pkeeling@interchange.ubc.ca

Affiliation Department of Botany, University of British Columbia, Vancouver, British Columbia, Canada

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: BAPW ALM PJK. Performed the experiments: BAPW CE LB ALM. Analyzed the data: BAPW ALM PJK. Contributed reagents/materials/analysis tools: YK KK ALM. Wrote the paper: BAPW ALM PJK.