About the Authors

David R. Maass
Institute of Environmental Science and Research Ltd., Porirua, Wellington, New Zealand

Gavin B. Harrison, Wayne R. Hein
Hopkirk Research Institute, AgResearch Grasslands Research Centre, Palmerston North, New Zealand

Warwick N. Grant
Genetics Department, La Trobe University, Melbourne, Australia

Charles B. Shoemaker
Department of Biomedical Sciences, Tufts Cummings School of Veterinary Medicine, North Grafton, Massachusetts, United States of America

Corresponding Author

Charles.Shoemaker@tufts.edu

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: DRM CBS. Performed the experiments: DRM. Analyzed the data: DRM GBH WNG WRH CBS. Contributed reagents/materials/analysis tools: GBH. Wrote the paper: DRM CBS.