About the Authors

Gareth J. Hughes, Samantha J. Lycett, Andrew Rambaut, Andrew J. Leigh Brown
Institute of Evolutionary Biology, School of Biological Sciences, University of Edinburgh, Edinburgh, Scotland, United Kingdom

Esther Fearnhill, David Dunn
Medical Research Council Clinical Trials Unit, London, United Kingdom

Corresponding Author

A.Leigh-Brown@ed.ac.uk

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: AJLB. Performed the experiments: GH. Analyzed the data: GH SJL. Contributed reagents/materials/analysis tools: EF DD AR. Wrote the paper: GH AJLB.