About the Authors

Gareth J. Hughes

Address Current address: Health Protection Agency East of England Regional Epidemiology Unit, Cambridge, United Kingdom

Affiliation Institute of Evolutionary Biology, School of Biological Sciences, University of Edinburgh, Edinburgh, Scotland, United Kingdom

Esther Fearnhill

Affiliation Medical Research Council Clinical Trials Unit, London, United Kingdom

David Dunn

Affiliation Medical Research Council Clinical Trials Unit, London, United Kingdom

Samantha J. Lycett

Affiliation Institute of Evolutionary Biology, School of Biological Sciences, University of Edinburgh, Edinburgh, Scotland, United Kingdom

Andrew Rambaut

Affiliation Institute of Evolutionary Biology, School of Biological Sciences, University of Edinburgh, Edinburgh, Scotland, United Kingdom

Andrew J. Leigh Brown

A.Leigh-Brown@ed.ac.uk

Affiliation Institute of Evolutionary Biology, School of Biological Sciences, University of Edinburgh, Edinburgh, Scotland, United Kingdom

on behalf of the UK HIV Drug Resistance Collaboration

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: AJLB. Performed the experiments: GH. Analyzed the data: GH SJL. Contributed reagents/materials/analysis tools: EF DD AR. Wrote the paper: GH AJLB.