About the Authors

Sally M. Williamson

Affiliation Department of Biology & Biochemistry, University of Bath, Bath, United Kingdom

Alan P. Robertson

Affiliation Department of Biomedical Sciences, Iowa State University, Ames, Iowa, United States of America

Laurence Brown

Current address: Department of Ophthalmology, University of Oxford, Oxford, United Kingdom

Affiliation MRC Functional Genomics Unit, Oxford, United Kingdom

Tracey Williams

Affiliation Veterinary Medicine Research & Development, Pfizer Animal Health, Kalamazoo, Michigan, United States of America

Debra J. Woods

Affiliation Veterinary Medicine Research & Development, Pfizer Animal Health, Kalamazoo, Michigan, United States of America

Richard J. Martin

Affiliation Department of Biomedical Sciences, Iowa State University, Ames, Iowa, United States of America

David B. Sattelle

Affiliation MRC Functional Genomics Unit, Oxford, United Kingdom

Adrian J. Wolstenholme

adrianw@uga.edu

Affiliation Department of Biology & Biochemistry, University of Bath, Bath, United Kingdom

Competing Interests

TW and DJW are employees of, and stockholders in, Pfizer Inc.

Author Contributions

Conceived and designed the experiments: SMW APR TW DJW RM DS AJW. Performed the experiments: SMW APR. Analyzed the data: SMW LB DS. Contributed reagents/materials/analysis tools: APR LB RM. Wrote the paper: SMW DJW RM DS AJW.