About the Authors

Soumitra Roy

Contributed equally to this work with: Soumitra Roy, Luk H. Vandenberghe

Affiliation Department of Pathology and Laboratory Medicine, University of Pennsylvania, Philadelphia, Pennsylvania, United States of America

Luk H. Vandenberghe

Contributed equally to this work with: Soumitra Roy, Luk H. Vandenberghe

Affiliation Department of Pathology and Laboratory Medicine, University of Pennsylvania, Philadelphia, Pennsylvania, United States of America

Sergey Kryazhimskiy

Affiliation Department of Biology, University of Pennsylvania, Philadelphia, Pennsylvania, United States of America

Rebecca Grant

Affiliation Department of Pathology and Laboratory Medicine, University of Pennsylvania, Philadelphia, Pennsylvania, United States of America

Roberto Calcedo

Affiliation Department of Pathology and Laboratory Medicine, University of Pennsylvania, Philadelphia, Pennsylvania, United States of America

Xin Yuan

Affiliation Department of Pathology and Laboratory Medicine, University of Pennsylvania, Philadelphia, Pennsylvania, United States of America

Martin Keough

Affiliation Department of Pathology and Laboratory Medicine, University of Pennsylvania, Philadelphia, Pennsylvania, United States of America

Arbans Sandhu

Affiliation Department of Pathology and Laboratory Medicine, University of Pennsylvania, Philadelphia, Pennsylvania, United States of America

Qiang Wang

Affiliation Vaccine Research Institute, Guangzhou, China

C. Angelica Medina-Jaszek

Affiliation Department of Pathology and Laboratory Medicine, University of Pennsylvania, Philadelphia, Pennsylvania, United States of America

Joshua B. Plotkin

Affiliation Department of Biology, University of Pennsylvania, Philadelphia, Pennsylvania, United States of America

James M. Wilson

wilsonjm@mail.med.upenn.edu

Affiliation Department of Pathology and Laboratory Medicine, University of Pennsylvania, Philadelphia, Pennsylvania, United States of America

Competing Interests

Soumitra Roy and Luk H. Vandenberghe are inventors on patents licensed to various companies. James M. Wilson is an inventor on patents licensed to various biopharmaceutical companies including ReGenX for which he has equity in, consults for and receives a grant from.

Author Contributions

Conceived and designed the experiments: SR LHV SK JBP JMW. Performed the experiments: SR LHV SK RC XY MK AS QW CAMJ JBP. Analyzed the data: SR LHV SK JBP JMW. Contributed reagents/materials/analysis tools: RG. Wrote the paper: SR LHV JBP JMW.