About the Authors

Reiko Yoshida

Affiliation Department of Global Epidemiology, Hokkaido University Research Center for Zoonosis Control, Sapporo, Hokkaido, Japan

Manabu Igarashi

Affiliation Department of Global Epidemiology, Hokkaido University Research Center for Zoonosis Control, Sapporo, Hokkaido, Japan

Hiroichi Ozaki

Affiliation Department of Veterinary Microbiology, Faculty of Agriculture, Tottori University, Tottori, Japan

Noriko Kishida

Affiliation Laboratory of Influenza Viruses, Department of Virology III, National Institute of Infectious Diseases, Musashimurayama, Tokyo, Japan

Daisuke Tomabechi

Affiliation Department of Global Epidemiology, Hokkaido University Research Center for Zoonosis Control, Sapporo, Hokkaido, Japan

Hiroshi Kida

Affiliations Department of Global Epidemiology, Hokkaido University Research Center for Zoonosis Control, Sapporo, Hokkaido, Japan, Laboratory of Microbiology, Department of Disease Control, Graduate School of Veterinary Medicine, Hokkaido University, Sapporo, Hokkaido, Japan

Kimihito Ito

Affiliation Department of Global Epidemiology, Hokkaido University Research Center for Zoonosis Control, Sapporo, Hokkaido, Japan

Ayato Takada

atakada@czc.hokudai.ac.jp

Affiliation Department of Global Epidemiology, Hokkaido University Research Center for Zoonosis Control, Sapporo, Hokkaido, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: RY AT. Performed the experiments: RY HO NK DT AT. Analyzed the data: RY MI HK KI AT. Wrote the paper: RY MI HO AT. Performed the molecular modeling: MI.