About the Authors

Joel W. Graff

Contributed equally to this work with: Joel W. Graff, Khalil Ettayebi

Affiliation Veterinary Molecular Biology, Montana State University, Bozeman, Montana, United States of America

Khalil Ettayebi

Contributed equally to this work with: Joel W. Graff, Khalil Ettayebi

Affiliation Veterinary Molecular Biology, Montana State University, Bozeman, Montana, United States of America

Michele E. Hardy

mhardy@montana.edu

Affiliation Veterinary Molecular Biology, Montana State University, Bozeman, Montana, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: JWG KE MEH. Performed the experiments: JWG KE. Analyzed the data: JWG KE MEH. Wrote the paper: JWG KE MEH.