About the Authors

Aleksandr S. Lipatov

Address Current address: Influenza Division, National Center for Immunization and Respiratory Diseases, Centers for Disease Control and Prevention, MS G-16, Atlanta, Georgia, United States of America

Affiliation Southeast Poultry Research Laboratory, Agricultural Research Service, United States Department of Agriculture, Athens, Georgia, United States of America

Yong Kuk Kwon

Affiliation Southeast Poultry Research Laboratory, Agricultural Research Service, United States Department of Agriculture, Athens, Georgia, United States of America

Luciana V. Sarmento

Affiliation Southeast Poultry Research Laboratory, Agricultural Research Service, United States Department of Agriculture, Athens, Georgia, United States of America

Kelly M. Lager

Affiliation National Animal Diseases Center, Agricultural Research Service, United States Department of Agriculture, Ames, Iowa, United States of America

Erica Spackman

Affiliation Southeast Poultry Research Laboratory, Agricultural Research Service, United States Department of Agriculture, Athens, Georgia, United States of America

David L. Suarez

Affiliation Southeast Poultry Research Laboratory, Agricultural Research Service, United States Department of Agriculture, Athens, Georgia, United States of America

David E. Swayne

david.swayne@ars.usda.gov

Affiliation Southeast Poultry Research Laboratory, Agricultural Research Service, United States Department of Agriculture, Athens, Georgia, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: AL YK KL DES. Performed the experiments: AL YK LS. Analyzed the data: AL YK LS KL ES DLS DES. Wrote the paper: AL DES. Performed sequencing and sequence analysis: ES DLS.