About the Authors

Eleanor M. Cottam

Affiliations Institute for Animal Health, Pirbright Laboratory, Pirbright, Woking, Surrey, United Kingdom, Division of Environmental and Evolutionary Biology, University of Glasgow, Glasgow, United Kingdom

Jemma Wadsworth

Affiliation Institute for Animal Health, Pirbright Laboratory, Pirbright, Woking, Surrey, United Kingdom

Andrew E. Shaw

Affiliation Institute for Animal Health, Pirbright Laboratory, Pirbright, Woking, Surrey, United Kingdom

Rebecca J. Rowlands

Affiliation Institute for Animal Health, Pirbright Laboratory, Pirbright, Woking, Surrey, United Kingdom

Lynnette Goatley

Affiliation Institute for Animal Health, Pirbright Laboratory, Pirbright, Woking, Surrey, United Kingdom

Sushila Maan

Affiliation Institute for Animal Health, Pirbright Laboratory, Pirbright, Woking, Surrey, United Kingdom

Narender S. Maan

Affiliation Institute for Animal Health, Pirbright Laboratory, Pirbright, Woking, Surrey, United Kingdom

Peter P. C. Mertens

Affiliation Institute for Animal Health, Pirbright Laboratory, Pirbright, Woking, Surrey, United Kingdom

Katja Ebert

Affiliation Institute for Animal Health, Pirbright Laboratory, Pirbright, Woking, Surrey, United Kingdom

Yanmin Li

Affiliation Institute for Animal Health, Pirbright Laboratory, Pirbright, Woking, Surrey, United Kingdom

Eoin D. Ryan

Affiliation Institute for Animal Health, Pirbright Laboratory, Pirbright, Woking, Surrey, United Kingdom

Nicholas Juleff

Affiliation Institute for Animal Health, Pirbright Laboratory, Pirbright, Woking, Surrey, United Kingdom

Nigel P. Ferris

Affiliation Institute for Animal Health, Pirbright Laboratory, Pirbright, Woking, Surrey, United Kingdom

John W. Wilesmith

Affiliation Animal Health and Welfare, Defra, London, United Kingdom

Daniel T. Haydon

Affiliation Division of Environmental and Evolutionary Biology, University of Glasgow, Glasgow, United Kingdom

Donald P. King

Affiliation Institute for Animal Health, Pirbright Laboratory, Pirbright, Woking, Surrey, United Kingdom

David J. Paton

Affiliation Institute for Animal Health, Pirbright Laboratory, Pirbright, Woking, Surrey, United Kingdom

Nick J. Knowles

nick.knowles@bbsrc.ac.uk

Affiliation Institute for Animal Health, Pirbright Laboratory, Pirbright, Woking, Surrey, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: EC DK DP NK. Performed the experiments: EC JW AS RR LG SM NM PM KE YL ER NJ NF. Analyzed the data: EC JWW DH DK DP NK. Contributed reagents/materials/analysis tools: EC YL ER NJ NF DH NK. Wrote the paper: EC DH DK DP NK.