About the Authors

Paul M Grandgenett

Affiliation Interdepartmental Genetics Program, University of Iowa, Iowa City, Iowa, United States of America

Keiko Otsu

Affiliation Department of Biochemistry, University of Iowa, Iowa City, Iowa, United States of America

Helen R Wilson

Affiliation Department of Biochemistry, University of Iowa, Iowa City, Iowa, United States of America

Mary E Wilson

Affiliations Department of Internal Medicine, University of Iowa, Iowa City, Iowa, United States of America , Department of Microbiology, University of Iowa, Iowa City, Iowa, United States of America , Veterans Affairs Medical Center, Iowa City, Iowa, United States of America

John E Donelson

To whom correspondence should be addressed. E-mail: john-donelson@uiowa.edu

Affiliations Interdepartmental Genetics Program, University of Iowa, Iowa City, Iowa, United States of America , Department of Biochemistry, University of Iowa, Iowa City, Iowa, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

PMG, MEW, and JED conceived and designed the experiments and wrote the paper. PMG, KO, and HRW performed the experiments. All authors analyzed the data.