About the Authors

Martha I Nelson

Affiliations Center for Infectious Disease Dynamics, The Pennsylvania State University, University Park, Pennsylvania, United States of America, Department of Biology, The Pennsylvania State University, University Park, Pennsylvania, United States of America

Lone Simonsen

Affiliation National Institute of Allergy and Infectious Diseases, National Institutes of Health, Bethesda, Maryland, United States of America

Cecile Viboud

Affiliation Fogarty International Center, National Institutes of Health, Bethesda, Maryland, United States of America

Mark A Miller

Affiliation Fogarty International Center, National Institutes of Health, Bethesda, Maryland, United States of America

Edward C Holmes

To whom correspondence should be addressed. E-mail: ech15@psu.edu

Affiliations Center for Infectious Disease Dynamics, The Pennsylvania State University, University Park, Pennsylvania, United States of America, Department of Biology, The Pennsylvania State University, University Park, Pennsylvania, United States of America, Fogarty International Center, National Institutes of Health, Bethesda, Maryland, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

MIN and ECH conceived the study. MIN performed the genome sequence analysis. MIN, LS, CV, MAM, and ECH wrote the paper.