About the Authors

Bryant McLaughlin

Affiliations Program in Microbial Pathogenesis and Host Defense, University of California San Francisco, San Francisco, California, United States of America , Department of Microbiology and Immunology, University of California San Francisco, San Francisco, California, United States of America

Janet S Chon

Affiliation Program in Microbial Pathogenesis and Host Defense, University of California San Francisco, San Francisco, California, United States of America

Jason A MacGurn

Affiliation Department of Microbiology and Immunology, University of California San Francisco, San Francisco, California, United States of America

Fredric Carlsson

¤ Current address: Department of Microbial Pathogenesis, Genentech, South San Francisco, California, United States of America

Affiliation Program in Microbial Pathogenesis and Host Defense, University of California San Francisco, San Francisco, California, United States of America

Terri L Cheng

Affiliation Program in Microbial Pathogenesis and Host Defense, University of California San Francisco, San Francisco, California, United States of America

Jeffery S Cox

Affiliation Department of Microbiology and Immunology, University of California San Francisco, San Francisco, California, United States of America

Eric J Brown

To whom correspondence should be addressed. E-mail: brown.eric@gene.com

¤ Current address: Department of Microbial Pathogenesis, Genentech, South San Francisco, California, United States of America

Affiliations Program in Microbial Pathogenesis and Host Defense, University of California San Francisco, San Francisco, California, United States of America , Department of Microbiology and Immunology, University of California San Francisco, San Francisco, California, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

BM and EJB conceived and designed the experiments and analyzed the data. BM, JSC, JAM, and TLC performed the experiments. BM, JAM, FC, JSC, and EJB contributed reagents/materials/analysis tools. BM, FC, JSC, and EJB wrote the paper.