About the Authors

John J Young

Contributed equally to this work with: John J Young, Jennifer L Bromberg-White

Affiliation Laboratory of Cancer and Developmental Cell Biology, Van Andel Research Institute, Grand Rapids, Michigan, United States of America

Jennifer L Bromberg-White

Contributed equally to this work with: John J Young, Jennifer L Bromberg-White

Affiliation Laboratory of Cancer and Developmental Cell Biology, Van Andel Research Institute, Grand Rapids, Michigan, United States of America

Cassandra Zylstra

Affiliation Laboratory of Cell Signaling and Carcinogenesis, Van Andel Research Institute, Grand Rapids, Michigan, United States of America

Joseph T Church

Affiliation Laboratory of Cancer and Developmental Cell Biology, Van Andel Research Institute, Grand Rapids, Michigan, United States of America

Elissa Boguslawski

Affiliation Laboratory of Cancer and Developmental Cell Biology, Van Andel Research Institute, Grand Rapids, Michigan, United States of America

James H Resau

Affiliation Laboratory of Analytical, Cellular, and Molecular Microscopy, Van Andel Research Institute, Grand Rapids, Michigan, United States of America

Bart O Williams

Affiliation Laboratory of Cell Signaling and Carcinogenesis, Van Andel Research Institute, Grand Rapids, Michigan, United States of America

Nicholas S Duesbery

To whom correspondence should be addressed. E-mail: duesbery@vai.org

Affiliation Laboratory of Cancer and Developmental Cell Biology, Van Andel Research Institute, Grand Rapids, Michigan, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

JJY, JLBW, BOW, and NSD conceived and designed the experiments and analyzed the data. JJY, JLBW, CZ, JTC, EB, JHR, and NSD performed the experiments. BOW contributed reagents/materials/analysis tools. NSD wrote the paper.