About the Authors

Cheryl D Chun

Contributed equally to this work with: Cheryl D Chun, Oliver W Liu

Affiliation Department of Biochemistry and Biophysics, University of California San Francisco, San Francisco, California, United States of America

Oliver W Liu

Contributed equally to this work with: Cheryl D Chun, Oliver W Liu

Affiliation Department of Biochemistry and Biophysics, University of California San Francisco, San Francisco, California, United States of America

Hiten D Madhani

To whom correspondence should be addressed. E-mail: hiten@biochem.ucsf.edu

Affiliation Department of Biochemistry and Biophysics, University of California San Francisco, San Francisco, California, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

CDC, OWL, and HDM conceived and designed the experiments, analyzed the data, and wrote the paper. CDC and OWL performed the experiments.