About the Authors

Kelly B McClellan

Affiliation Department of Pathology and Immunology, Washington University School of Medicine, Saint Louis, Missouri, United States of America

Shivaprakash Gangappa

Affiliation Emory Transplant Center, Emory University School of Medicine, Atlanta, Georgia, United States of America

Samuel H Speck

Affiliation Division of Microbiology and Immunology, Yerkes Regional Primate Research Center, Emory University, Atlanta, Georgia, United States of America

Herbert W. Virgin IV

To whom correspondence should be addressed. E-mail: virgin@wustl.edu

Affiliation Department of Pathology and Immunology, Washington University School of Medicine, Saint Louis, Missouri, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

KBM, SG, SHS, and HWV conceived and designed the experiments. KBM and SG performed the experiments. KBM, SG, and HWV analyzed the data. KBM and HWV wrote the paper.