About the Authors

Kenneth A. Field

kfield@bucknell.edu

Affiliation Department of Biology, Bucknell University, Lewisburg, Pennsylvania, United States of America

Joseph S. Johnson

Affiliation Department of Biology, Bucknell University, Lewisburg, Pennsylvania, United States of America

Thomas M. Lilley

Affiliation Department of Biology, Bucknell University, Lewisburg, Pennsylvania, United States of America

Sophia M. Reeder

Affiliation Department of Biology, Bucknell University, Lewisburg, Pennsylvania, United States of America

Elizabeth J. Rogers

Affiliation Department of Biology, Bucknell University, Lewisburg, Pennsylvania, United States of America

Melissa J. Behr

Affiliation Department of Pathobiological Sciences, School of Veterinary Medicine, University of Wisconsin-Madison, Madison, Wisconsin, United States of America

DeeAnn M. Reeder

Affiliation Department of Biology, Bucknell University, Lewisburg, Pennsylvania, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: KAF JSJ DMR. Performed the experiments: KAF SMR EJR MJB. Analyzed the data: KAF. Wrote the paper: KAF JSJ TML DMR.