About the Authors

Chris Coward

cc122@cam.ac.uk

Affiliation University of Cambridge, Department of Veterinary Medicine, Cambridge, United Kingdom

Olivier Restif

Affiliation University of Cambridge, Department of Veterinary Medicine, Cambridge, United Kingdom

Richard Dybowski

Affiliation University of Cambridge, Department of Veterinary Medicine, Cambridge, United Kingdom

Andrew J. Grant

Affiliation University of Cambridge, Department of Veterinary Medicine, Cambridge, United Kingdom

Duncan J. Maskell

Affiliation University of Cambridge, Department of Veterinary Medicine, Cambridge, United Kingdom

Pietro Mastroeni

Affiliation University of Cambridge, Department of Veterinary Medicine, Cambridge, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: CC OR AJG PM. Performed the experiments: CC. Analyzed the data: CC OR. Contributed reagents/materials/analysis tools: AJG OR. Contributed to the writing of the manuscript: CC OR RD AJG DJM PM. Provided advice on statistical analysis: RD.