About the Authors

Samuel K. Buxton

Contributed equally to this work with: Samuel K. Buxton, Claude L. Charvet

Affiliations Department of Biomedical Sciences, College of Veterinary Medicine, Iowa State University, Ames, Iowa, United States of America, INRA, UR1282 Infectiologie Animale et Santé Publique, Nouzilly, France

Claude L. Charvet

Contributed equally to this work with: Samuel K. Buxton, Claude L. Charvet

Affiliations INRA, UR1282 Infectiologie Animale et Santé Publique, Nouzilly, France, Université François Rabelais de Tours, UMR1282 Infectiologie et Santé Publique, Tours, France

Cedric Neveu

Affiliations INRA, UR1282 Infectiologie Animale et Santé Publique, Nouzilly, France, Université François Rabelais de Tours, UMR1282 Infectiologie et Santé Publique, Tours, France

Jacques Cabaret

Affiliations INRA, UR1282 Infectiologie Animale et Santé Publique, Nouzilly, France, Université François Rabelais de Tours, UMR1282 Infectiologie et Santé Publique, Tours, France

Jacques Cortet

Affiliations INRA, UR1282 Infectiologie Animale et Santé Publique, Nouzilly, France, Université François Rabelais de Tours, UMR1282 Infectiologie et Santé Publique, Tours, France

Nicolas Peineau

Affiliation Université François Rabelais de Tours, Département de Physiologie Animale, Tours, France

Melanie Abongwa

Affiliation Department of Biomedical Sciences, College of Veterinary Medicine, Iowa State University, Ames, Iowa, United States of America

Elise Courtot

Affiliation INRA, UR1282 Infectiologie Animale et Santé Publique, Nouzilly, France

Alan P. Robertson

Affiliation Department of Biomedical Sciences, College of Veterinary Medicine, Iowa State University, Ames, Iowa, United States of America

Richard J. Martin

rjmartin@iastate.edu

Affiliation Department of Biomedical Sciences, College of Veterinary Medicine, Iowa State University, Ames, Iowa, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: SKB CLC CN APR RJM. Performed the experiments: SKB CLC JCo MA APR. Analyzed the data: SKB CLC CN APR RJM. Contributed reagents/materials/analysis tools: CLC CN NP APR RJM EC. Wrote the paper: SKB CLC CN JCa APR NP RJM.