About the Authors

William G. C. Horsnell

Contributed equally to this work with: William G. C. Horsnell, Matthew G. Darby

Affiliation International Centre for Genetic Engineering and Biotechnology (ICGEB) and Institute of Infectious Disease and Molecular Medicine (IIDMM), Division of Immunology, University of Cape Town, Cape Town, South Africa

Matthew G. Darby

Contributed equally to this work with: William G. C. Horsnell, Matthew G. Darby

Affiliation International Centre for Genetic Engineering and Biotechnology (ICGEB) and Institute of Infectious Disease and Molecular Medicine (IIDMM), Division of Immunology, University of Cape Town, Cape Town, South Africa

Jennifer C. Hoving

Affiliation International Centre for Genetic Engineering and Biotechnology (ICGEB) and Institute of Infectious Disease and Molecular Medicine (IIDMM), Division of Immunology, University of Cape Town, Cape Town, South Africa

Natalie Nieuwenhuizen

Affiliation International Centre for Genetic Engineering and Biotechnology (ICGEB) and Institute of Infectious Disease and Molecular Medicine (IIDMM), Division of Immunology, University of Cape Town, Cape Town, South Africa

Henry J. McSorley

Affiliation Institute of Immunology and Infection Research, University of Edinburgh, Ashworth Laboratories, Edinburgh, United Kingdom

Hlumani Ndlovu

Affiliation International Centre for Genetic Engineering and Biotechnology (ICGEB) and Institute of Infectious Disease and Molecular Medicine (IIDMM), Division of Immunology, University of Cape Town, Cape Town, South Africa

Saeeda Bobat

Affiliation School of Immunity and Infection and Medical Research Council Centre for Immune Regulation, University of Birmingham, Birmingham, United Kingdom

Matti Kimberg

Affiliation International Centre for Genetic Engineering and Biotechnology (ICGEB) and Institute of Infectious Disease and Molecular Medicine (IIDMM), Division of Immunology, University of Cape Town, Cape Town, South Africa

Frank Kirstein

Affiliation International Centre for Genetic Engineering and Biotechnology (ICGEB) and Institute of Infectious Disease and Molecular Medicine (IIDMM), Division of Immunology, University of Cape Town, Cape Town, South Africa

Anthony J. Cutler

Current address: Histocompatibility and Immunogenetics, NHSBT, Colindale Centre, London, United Kingdom.

Affiliation International Centre for Genetic Engineering and Biotechnology (ICGEB) and Institute of Infectious Disease and Molecular Medicine (IIDMM), Division of Immunology, University of Cape Town, Cape Town, South Africa

Benjamin DeWals

Current address: Immunology-Vaccinology, Department of Infectious and Parasitic Diseases, Faculty of Veterinary Medicine, University of Liège, Liège, Belgium.

Affiliation International Centre for Genetic Engineering and Biotechnology (ICGEB) and Institute of Infectious Disease and Molecular Medicine (IIDMM), Division of Immunology, University of Cape Town, Cape Town, South Africa

Adam F. Cunningham

Affiliation School of Immunity and Infection and Medical Research Council Centre for Immune Regulation, University of Birmingham, Birmingham, United Kingdom

Frank Brombacher

fbrombac@mweb.co.za

Affiliation International Centre for Genetic Engineering and Biotechnology (ICGEB) and Institute of Infectious Disease and Molecular Medicine (IIDMM), Division of Immunology, University of Cape Town, Cape Town, South Africa

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: WGCH MGD FB AJC. Performed the experiments: WGCH MGD JCH NN HN SB MK FK BD AFC HJM. Analyzed the data: WGCH MGD JCH NN. Contributed reagents/materials/analysis tools: WGCH FB AJC. Wrote the paper: WGCH MGD FB AJC.