About the Authors

Adam L. Vanarsdall

Affiliation Department of Molecular Microbiology and Immunology, Oregon Health and Science University, Portland, Oregon, United States of America

Todd W. Wisner

Affiliation Department of Molecular Microbiology and Immunology, Oregon Health and Science University, Portland, Oregon, United States of America

Hetian Lei

Affiliation The Schepens Eye Research Institute, Department of Ophthalmology, Harvard Medical School, Boston, Massachusetts, United States of America

Andrius Kazlauskas

Affiliation The Schepens Eye Research Institute, Department of Ophthalmology, Harvard Medical School, Boston, Massachusetts, United States of America

David C. Johnson

johnsoda@ohsu.edu

Affiliation Department of Molecular Microbiology and Immunology, Oregon Health and Science University, Portland, Oregon, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: ALV DCJ. Performed the experiments: ALV TWW. Analyzed the data: ALV TWW DCJ. Contributed reagents/materials/analysis tools: HL AK. Wrote the paper: DCJ ALV. Read and aided in revising the paper: HL AK TWW.