About the Authors

Shannon Dennis

Affiliations Fred Hutchinson Cancer Research Center, Seattle, Washington, United States of America, Howard Hughes Medical Institute, Seattle, Washington, United States of America, Molecular and Cellular Biology Program, University of Washington, Seattle, Washington, United States of America

Ujwal Sheth

Contributed equally to this work with: Ujwal Sheth, Jessica L. Feldman

Affiliations Fred Hutchinson Cancer Research Center, Seattle, Washington, United States of America, Howard Hughes Medical Institute, Seattle, Washington, United States of America

Jessica L. Feldman

Contributed equally to this work with: Ujwal Sheth, Jessica L. Feldman

Affiliations Fred Hutchinson Cancer Research Center, Seattle, Washington, United States of America, Howard Hughes Medical Institute, Seattle, Washington, United States of America

Kathryn A. English

Affiliations Fred Hutchinson Cancer Research Center, Seattle, Washington, United States of America, Howard Hughes Medical Institute, Seattle, Washington, United States of America

James R. Priess

jpriess@fhcrc.org

Affiliations Fred Hutchinson Cancer Research Center, Seattle, Washington, United States of America, Howard Hughes Medical Institute, Seattle, Washington, United States of America, Molecular and Cellular Biology Program, University of Washington, Seattle, Washington, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: SD US JLF JRP. Performed the experiments: SD US JLF KAE JRP. Analyzed the data: SD US JLF KAE JRP. Wrote the paper: SD US JLF JRP.