About the Authors

Matthew B. Frieman

Contributed equally to this work with: Matthew B. Frieman, Jun Chen

Current address: Department of Microbiology and Immunology, University of Maryland School of Medicine, Baltimore, Maryland, United States of America

Affiliation Department of Epidemiology, University of North Carolina at Chapel Hill, Chapel Hill, North Carolina, United States of America

Jun Chen

Contributed equally to this work with: Matthew B. Frieman, Jun Chen

Current address: Laboratory of Immunology, National Eye Institute, NIH, Bethesda, Maryland, United States of America

Affiliation Laboratory of Infectious Diseases, NIAID, NIH, Bethesda, Maryland, United States of America

Thomas E. Morrison

Current address: Department of Microbiology, University of Colorado Denver, Aurora, Colorado, United States of America

Affiliation Department of Microbiology and Immunology, University of North Carolina at Chapel Hill, Chapel Hill, North Carolina, United States of America

Alan Whitmore

Affiliation Department of Microbiology and Immunology, University of North Carolina at Chapel Hill, Chapel Hill, North Carolina, United States of America

William Funkhouser

Affiliation Department of Anatomic Pathology and Surgical Pathology, University of North Carolina at Chapel Hill, Chapel Hill, North Carolina, United States of America

Jerrold M. Ward

Affiliations Comparative Medicine Branch, NIAID, NIH, Bethesda, Maryland, United States of America, Laboratory of Immunopathology, NIAID, NIH, Bethesda, Maryland, United States of America

Elaine W. Lamirande

Affiliation Laboratory of Infectious Diseases, NIAID, NIH, Bethesda, Maryland, United States of America

Anjeanette Roberts

Affiliation Laboratory of Infectious Diseases, NIAID, NIH, Bethesda, Maryland, United States of America

Mark Heise

Affiliation Department of Microbiology and Immunology, University of North Carolina at Chapel Hill, Chapel Hill, North Carolina, United States of America

Kanta Subbarao

Affiliation Laboratory of Infectious Diseases, NIAID, NIH, Bethesda, Maryland, United States of America

Ralph S. Baric

ralph_baric@unc.edu

Affiliations Department of Epidemiology, University of North Carolina at Chapel Hill, Chapel Hill, North Carolina, United States of America, Department of Microbiology and Immunology, University of North Carolina at Chapel Hill, Chapel Hill, North Carolina, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: MBF JC KS RSB. Performed the experiments: MBF JC TEM AW EWL AR. Analyzed the data: MBF JC TEM AW WF JMW EWL AR MH KS. Contributed reagents/materials/analysis tools: MBF JC TEM AW MH KS. Wrote the paper: MBF JC.