About the Authors

M. Khyrul Islam

Affiliation Laboratory of Parasitic Diseases, National Institute of Animal Health, National Agricultural and Food Research Organization, Tsukuba, Ibaraki, Japan

Naotoshi Tsuji

tsujin@affrc.go.jp

Affiliation Laboratory of Parasitic Diseases, National Institute of Animal Health, National Agricultural and Food Research Organization, Tsukuba, Ibaraki, Japan

Takeharu Miyoshi

Affiliation Laboratory of Parasitic Diseases, National Institute of Animal Health, National Agricultural and Food Research Organization, Tsukuba, Ibaraki, Japan

M. Abdul Alim

Affiliation Laboratory of Parasitic Diseases, National Institute of Animal Health, National Agricultural and Food Research Organization, Tsukuba, Ibaraki, Japan

Xiaohong Huang

Affiliation Laboratory of Parasitic Diseases, National Institute of Animal Health, National Agricultural and Food Research Organization, Tsukuba, Ibaraki, Japan

Takeshi Hatta

Affiliation Laboratory of Parasitic Diseases, National Institute of Animal Health, National Agricultural and Food Research Organization, Tsukuba, Ibaraki, Japan

Kozo Fujisaki

Affiliation Department of Emerging Infectious Diseases, School of Veterinary Medicine, University of Kagoshima, Kagoshima, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: MKI NT KF. Performed the experiments: MKI NT TM MAA XH TH. Analyzed the data: MKI NT KF. Wrote the paper: MKI NT KF.