About the Authors

Robert T. Wheeler

robert.wheeler@umit.maine.edu

Current address: University of Maine, BMMB Department, Orono, Maine, United States of America

Affiliation Whitehead Institute for Biomedical Research, Cambridge, Massachusetts, United States of America

Diana Kombe

Affiliation Whitehead Institute for Biomedical Research, Cambridge, Massachusetts, United States of America

Sudeep D. Agarwala

Affiliation Whitehead Institute for Biomedical Research, Cambridge, Massachusetts, United States of America

Gerald R. Fink

Affiliation Whitehead Institute for Biomedical Research, Cambridge, Massachusetts, United States of America

Competing Interests

This work was funded in part by a grant from Merck, Inc. to GRF and RTW. This sponsor had no role in the study or manuscript beyond funding the work.

Author Contributions

Conceived and designed the experiments: RTW GRF. Performed the experiments: RTW DK. Analyzed the data: RTW. Contributed reagents/materials/analysis tools: SDA. Wrote the paper: RTW GRF.